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Paul Talalay and researchers at Johns Hopkins University linked these findings (Prochaska et al., 1992) and identified sulforaphane, isolated from broccoli, as a potent inducer of phase II detoxification enzymes including glutathione-S-transferase (GST) and NAD(P)H oxidoreductase 1 (NQO1) (Zhang et al., 1992)

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Therefore, although GSH is potentially able to reduce non-native disulphide bonds in newly synthesized proteins, there is a kinetic advantage for this reduction to be catalysed by the ER oxidoreductases